.htaccess is a configuration file for used on web servers such as the Apache Web Server software. The .htaccess file can be used to alter or change or override the server configuration settings of the Apache Web Server software. It is used to make configuration changes on a per-directory basis. It can be used to add additional functionality to the Apache Web Server.
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WordPress will always flush the Rewrite URL's or permalinks whenever you update your permalink structure. simply update your permalinks. Read More